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Govindaraju, G S and Jayasankar, P (2004) Taxonomic Relationship Among Seven Species of Groupers (Genus Epinephelus; Family Serranidae) as Revealed by RAPD Fingerprinting. Marine Biotechnology, 6. pp. 229-237.
Govindaraju, G S (2002) Genetic profiling of groupers of the Genus Epinephelus (Family: Serranidae) using DNA- Level markers. Masters thesis, Central Institute of Fisheries Education.